:root {
  --ink: #17243a;
  --ink-deep: #0f1928;
  --ink-soft: #263750;
  --paper: #f1f5f4;
  --surface: #ffffff;
  --line: #d7e0de;
  --muted: #5d6672;
  --teal: #287a76;
  --teal-dark: #195d5a;
  --crimson: #7d3340;
  --max: 1160px;
  --radius: 12px;
  --shadow: 0 14px 36px rgba(23, 36, 58, .08);
}

* { box-sizing: border-box; }
html { min-height: 100%; scroll-behavior: smooth; }
body {
  min-height: 100vh;
  min-height: 100svh;
  display: flex;
  flex-direction: column;
  margin: 0;
  color: var(--ink);
  background: var(--paper);
  font-family: Inter, ui-sans-serif, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  font-size: 16px;
  line-height: 1.68;
  -webkit-font-smoothing: antialiased;
}
body > main { flex: 1 0 auto; }
body > [data-site-header], body > [data-site-footer] { flex: 0 0 auto; }
a { color: inherit; }
button, input, select, textarea { font: inherit; }
button, a { -webkit-tap-highlight-color: transparent; }
h1, h2, h3, h4, p { margin-top: 0; }
h1, h2, h3 {
  color: var(--ink);
  font-family: Inter, ui-sans-serif, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  font-weight: 760;
  letter-spacing: -.045em;
  line-height: 1.08;
}
.skip-link { position: fixed; left: 1rem; top: -5rem; z-index: 100; padding: .7rem 1rem; background: white; color: var(--ink); border: 1px solid var(--line); border-radius: 4px; }
.skip-link:focus { top: 1rem; }
.shell { width: min(calc(100% - 40px), var(--max)); margin-inline: auto; }

.site-header {
  position: sticky;
  top: 0;
  z-index: 50;
  background: rgba(255, 253, 248, .96);
  border-bottom: 1px solid var(--line);
  backdrop-filter: blur(14px);
}
.nav-wrap { min-height: 78px; display: flex; align-items: center; justify-content: space-between; gap: 2rem; }
.brand { display: inline-flex; align-items: center; gap: .8rem; color: var(--ink); text-decoration: none; }
.brand-mark { width: 40px; height: 36px; display: grid; place-items: center; background: var(--ink); color: white; border-radius: 3px; font-size: .58rem; font-weight: 800; letter-spacing: .08em; }
.brand-text { display: grid; line-height: 1.12; }
.brand-text strong { font-size: 1.02rem; font-weight: 800; letter-spacing: -.02em; }
.brand-text small { margin-top: .2rem; color: var(--muted); font-size: .64rem; letter-spacing: .09em; text-transform: uppercase; }
.site-nav { display: flex; align-items: center; gap: 1.35rem; }
.site-nav a { position: relative; color: #4d5867; text-decoration: none; font-size: .86rem; font-weight: 600; padding: 1.65rem 0 1.45rem; }
.site-nav a::after { content: ""; position: absolute; left: 0; right: 0; bottom: 1rem; height: 2px; background: transparent; }
.site-nav a:hover { color: var(--teal-dark); }
.site-nav a[aria-current="page"] { color: var(--ink); }
.site-nav a[aria-current="page"]::after { background: var(--teal); }
.nav-toggle { display: none; border: 1px solid var(--line); color: var(--ink); background: var(--surface); border-radius: 4px; width: 42px; height: 42px; }

.hero {
  position: relative;
  overflow: hidden;
  color: white;
  background: var(--ink-deep);
  border-bottom: 1px solid rgba(255,255,255,.1);
}
.hero::before {
  content: "";
  position: absolute;
  inset: 0;
  background: radial-gradient(circle at 75% 35%, rgba(40,122,118,.19), transparent 34%), radial-gradient(circle at 30% 85%, rgba(125,51,64,.12), transparent 28%);
  pointer-events: none;
}
.hero-grid { position: relative; display: grid; grid-template-columns: 1.04fr .96fr; gap: clamp(3rem, 7vw, 6rem); align-items: center; min-height: 690px; padding: 6.5rem 0 7rem; }
.hero-copy { position: relative; }
.hero-copy::before { content: ""; position: absolute; left: -24px; top: 2px; width: 3px; height: 54px; background: #59b7b1; }
.eyebrow { margin: 0 0 1rem; color: var(--teal-dark); font-size: .7rem; font-weight: 800; letter-spacing: .14em; line-height: 1.3; text-transform: uppercase; }
.hero .eyebrow { color: #78c7c2; }
.hero h1 { margin-bottom: 1.6rem; max-width: 12ch; color: white; font-size: clamp(3.2rem, 6.4vw, 5.8rem); }
.hero h1 span { color: #65c4be; }
.hero-lede { max-width: 690px; color: #c0cad5; font-size: clamp(1.05rem, 1.6vw, 1.25rem); }
.hero-actions { display: flex; flex-wrap: wrap; gap: .9rem; margin-top: 2rem; }
.button { display: inline-flex; align-items: center; justify-content: center; min-height: 46px; padding: .65rem 1.05rem; border: 1px solid transparent; border-radius: 4px; font-size: .84rem; font-weight: 750; text-decoration: none; transition: background .18s ease, color .18s ease, border-color .18s ease; }
.button-primary { color: var(--ink-deep); background: #65c4be; }
.button-dark { color: white; background: var(--ink); }
.button-primary:hover, .button-dark:hover { background: var(--teal-dark); }
.hero .button-primary:hover { color: white; background: var(--teal); }
.button-secondary { color: var(--ink); border-color: var(--line); background: transparent; }
.hero .button-secondary { color: white; border-color: rgba(255,255,255,.28); }
.button-secondary:hover, .button-ghost:hover { border-color: var(--teal); color: var(--teal-dark); }
.button-ghost { border-color: var(--line); background: var(--surface); }

.research-visual { position: relative; min-height: 470px; overflow: hidden; border: 1px solid rgba(255,255,255,.16); border-radius: 16px; background: rgba(255,255,255,.035); box-shadow: 0 28px 80px rgba(0,0,0,.22); backdrop-filter: blur(8px); }
.visual-heading { position: absolute; z-index: 2; left: 1.4rem; right: 1.4rem; top: 1.2rem; display: flex; justify-content: space-between; gap: 1rem; color: #91a1b2; font-size: .65rem; letter-spacing: .12em; text-transform: uppercase; }
.visual-heading strong { color: white; font-weight: 750; letter-spacing: .02em; text-transform: none; }
#research-canvas { position: absolute; inset: 48px 0 52px; width: 100%; height: calc(100% - 100px); }
.visual-legend { position: absolute; z-index: 2; left: 1.4rem; right: 1.4rem; bottom: 1.1rem; display: flex; flex-wrap: wrap; gap: 1rem; color: #9facba; font-size: .64rem; }
.visual-legend span { display: inline-flex; align-items: center; gap: .4rem; }
.visual-legend i { width: 7px; height: 7px; border-radius: 50%; }
.legend-molecular { background: #65c4be; }
.legend-clinical { background: #d58996; }
.legend-translation { background: #f0d78a; }

.home-news { padding: 3.4rem 0 4rem; background: var(--surface); border-bottom: 1px solid var(--line); }
.home-news-heading { display: flex; align-items: end; justify-content: space-between; gap: 2rem; margin-bottom: 1.5rem; }
.home-news-heading .eyebrow { margin-bottom: .45rem; }
.home-news-heading h2 { margin-bottom: 0; font-size: clamp(1.7rem, 3vw, 2.4rem); }
.home-news-heading > a { color: var(--teal-dark); font-size: .76rem; font-weight: 800; text-decoration: none; }
.news-strip { display: grid; grid-template-columns: repeat(3, 1fr); border: 1px solid var(--line); }
.news-brief { display: grid; grid-template-columns: 46px 1fr auto; gap: .95rem; align-items: start; min-height: 122px; padding: 1.15rem; border-right: 1px solid var(--line); color: var(--ink); text-decoration: none; transition: background .18s ease; }
.news-brief:last-child { border-right: 0; }
.news-brief:hover { background: var(--paper); }
.news-brief time { color: var(--crimson); font-size: .72rem; font-weight: 800; letter-spacing: .05em; }
.news-brief span { display: block; margin-bottom: .38rem; color: var(--teal-dark); font-size: .68rem; font-weight: 800; letter-spacing: .09em; text-transform: uppercase; }
.news-brief h3 { margin: 0; font-size: 1rem; line-height: 1.42; letter-spacing: -.015em; }
.news-brief b { color: var(--teal-dark); font-size: 1rem; }

.stats-band { color: white; background: var(--teal-dark); }
.stats-grid { min-height: 118px; display: grid; grid-template-columns: .65fr .65fr .9fr 1.45fr auto; align-items: center; gap: 1.5rem; }
.stats-grid > div { padding-right: 1rem; border-right: 1px solid rgba(255,255,255,.2); }
.stats-grid strong, .stats-grid span { display: block; }
.stats-grid strong { font-size: 1.75rem; line-height: 1; }
.stats-grid span { margin-top: .4rem; color: #b8d9d6; font-size: .65rem; letter-spacing: .08em; text-transform: uppercase; }
.stats-grid p { margin: 0; color: #d4e6e4; font-size: .82rem; }
.stats-grid a { color: white; font-size: .78rem; font-weight: 800; text-decoration: none; white-space: nowrap; }

.section { padding: 6.5rem 0; }
.section-head { display: grid; grid-template-columns: .85fr 1.15fr; gap: 4rem; align-items: end; margin-bottom: 3rem; }
.section-head h2, .page-hero h1 { margin-bottom: 0; font-size: clamp(2.5rem, 4.7vw, 4.4rem); }
.section-intro { margin: 0; color: var(--muted); font-size: 1.04rem; }
.mission-grid { display: grid; grid-template-columns: 1.05fr .95fr; gap: 1.25rem; }
.mission-copy { padding: clamp(2rem, 4vw, 3.6rem); background: var(--surface); border: 1px solid var(--line); border-left: 4px solid var(--teal); }
.mission-copy h2 { font-size: clamp(2.1rem, 3.5vw, 3.35rem); }
.mission-copy p { color: var(--muted); font-size: 1.05rem; }
.metric-grid { display: grid; grid-template-columns: 1fr 1fr; gap: 1rem; }
.metric { min-height: 146px; padding: 1.35rem; background: var(--surface); border: 1px solid var(--line); border-top: 3px solid var(--teal); display: flex; flex-direction: column; justify-content: space-between; }
.metric strong { color: var(--ink); font-size: 1.35rem; font-weight: 800; }
.metric span { color: var(--muted); font-size: .8rem; }

.dark-section { color: white; background: var(--ink); }
.dark-section h2, .dark-section h3 { color: white; }
.dark-section .eyebrow { color: #8fc9c5; }
.dark-section .section-intro { color: #c0c8d2; }
.research-explorer { display: grid; grid-template-columns: .78fr 1.22fr; min-height: 480px; border: 1px solid rgba(255,255,255,.16); border-radius: 12px; overflow: hidden; background: rgba(255,255,255,.03); }
.theme-tabs { display: grid; align-content: stretch; border-right: 1px solid rgba(255,255,255,.16); }
.theme-tab { display: grid; grid-template-columns: 34px 1fr; grid-template-rows: auto auto; column-gap: .75rem; padding: 1.5rem; color: #b8c4cf; background: transparent; border: 0; border-bottom: 1px solid rgba(255,255,255,.13); text-align: left; cursor: pointer; transition: background .2s ease, color .2s ease; }
.theme-tab:last-child { border-bottom: 0; }
.theme-tab:hover { color: white; background: rgba(255,255,255,.045); }
.theme-tab[aria-selected="true"] { color: white; background: rgba(101,196,190,.12); box-shadow: inset 3px 0 #65c4be; }
.theme-tab > span { grid-row: 1 / span 2; color: #65c4be; font-size: .7rem; font-weight: 800; letter-spacing: .08em; }
.theme-tab strong { font-size: .92rem; }
.theme-tab small { margin-top: .25rem; color: #8f9eae; font-size: .7rem; }
.theme-detail { position: relative; padding: clamp(2rem, 5vw, 4rem); outline: none; }
.theme-detail::after { content: ""; position: absolute; right: -60px; bottom: -80px; width: 260px; height: 260px; border: 1px solid rgba(101,196,190,.15); border-radius: 50%; box-shadow: 0 0 0 46px rgba(101,196,190,.035), 0 0 0 92px rgba(101,196,190,.025); pointer-events: none; }
.theme-detail h3 { position: relative; z-index: 1; font-size: clamp(2rem, 4vw, 3.4rem); }
.theme-detail > p:not(.eyebrow) { position: relative; z-index: 1; max-width: 680px; color: #c0cad5; font-size: 1rem; }
.evidence-grid { position: relative; z-index: 1; display: grid; grid-template-columns: repeat(3, 1fr); gap: 1px; margin-top: 2.2rem; background: rgba(255,255,255,.13); }
.evidence-grid > div { min-height: 132px; padding: 1rem; background: var(--ink); }
.evidence-grid span, .evidence-grid strong { display: block; }
.evidence-grid span { margin-bottom: .8rem; color: #65c4be; font-size: .62rem; font-weight: 800; letter-spacing: .1em; text-transform: uppercase; }
.evidence-grid strong { color: #e0e6ec; font-size: .76rem; line-height: 1.55; }
.topic-keywords { position: relative; z-index: 1; display: flex; flex-wrap: wrap; gap: .45rem; margin-top: 1.4rem; }
.topic-keywords span { padding: .3rem .55rem; color: #d4dde5; border: 1px solid rgba(255,255,255,.16); border-radius: 999px; font-size: .66rem; }

.pipeline { display: grid; grid-template-columns: repeat(4, 1fr); gap: 0; border: 1px solid var(--line); background: var(--surface); }
.pipeline-step { min-height: 220px; padding: 1.6rem; background: var(--surface); border-right: 1px solid var(--line); }
.pipeline-step:last-child { border-right: 0; }
.pipeline-step b { display: block; margin-bottom: 3rem; color: var(--crimson); font-size: .7rem; letter-spacing: .08em; text-transform: uppercase; }
.pipeline-step h3 { font-size: 1.35rem; }
.pipeline-step p { color: var(--muted); font-size: .87rem; }

.cards { display: grid; grid-template-columns: repeat(3, 1fr); gap: 1rem; }
.card { padding: 1.55rem; border: 1px solid var(--line); border-radius: 6px; background: var(--surface); }
.card:hover { border-color: #a7aaa3; }
.card-meta { display: flex; justify-content: space-between; gap: 1rem; margin-bottom: 1.2rem; color: var(--teal-dark); font-size: .66rem; font-weight: 800; letter-spacing: .08em; text-transform: uppercase; }
.card h3 { font-size: 1.28rem; }
.card p { color: var(--muted); font-size: .88rem; }
.card-link { color: var(--ink); font-weight: 750; font-size: .8rem; text-decoration-thickness: 1px; text-underline-offset: 4px; }
.section-cta { margin-top: 2rem; display: flex; flex-wrap: wrap; justify-content: flex-end; gap: .75rem; }

.page-hero { padding: 6.5rem 0 4.5rem; color: white; background: var(--ink); border-bottom: 5px solid var(--teal); }
.page-hero .shell { display: grid; grid-template-columns: 1.15fr .85fr; gap: 4rem; align-items: end; }
.page-hero h1 { color: white; }
.page-hero .eyebrow { color: #9fd0cd; }
.page-hero p { margin: 0; color: #c3ccd5; font-size: 1.02rem; }
.publication-intro p { margin-bottom: 1.25rem; }
.publication-links { display: flex; flex-wrap: wrap; gap: .6rem; }
.publication-links .button { min-height: 40px; color: white; border-color: rgba(255,255,255,.28); background: rgba(255,255,255,.06); }
.publication-links .button:hover { border-color: #9fd0cd; background: rgba(101,196,190,.13); }
.filters { display: grid; grid-template-columns: 1.5fr .7fr .8fr; gap: .8rem; margin-bottom: 1.5rem; }
.field { width: 100%; min-height: 48px; padding: .7rem .9rem; color: var(--ink); background: var(--surface); border: 1px solid var(--line); border-radius: 4px; }
.field:focus { outline: 2px solid rgba(40,122,118,.22); border-color: var(--teal); }
.result-count { color: var(--muted); font-size: .82rem; margin-bottom: 1rem; }
.publication-list { display: grid; gap: .65rem; }
.publication { display: grid; grid-template-columns: 90px 1fr; gap: 1.5rem; padding: 1.35rem; border: 1px solid var(--line); border-radius: 4px; background: var(--surface); }
.publication-year { color: var(--teal-dark); font-size: 1.08rem; font-weight: 800; }
.publication h2 { margin-bottom: .55rem; font-family: inherit; font-size: 1.05rem; font-weight: 750; letter-spacing: -.01em; line-height: 1.35; }
.publication .authors { margin-bottom: .35rem; color: var(--muted); font-size: .82rem; }
.publication .journal { margin-bottom: .55rem; color: var(--teal-dark); font-family: Georgia, "Times New Roman", serif; font-size: .96rem; font-style: italic; font-weight: 700; letter-spacing: .01em; line-height: 1.3; }
.publication .me { color: var(--crimson); font-weight: 800; }
.empty-state { padding: 4rem; text-align: center; border: 1px dashed #aaa; background: var(--surface); }

.people-roster { background: var(--paper); }
.people-directory { border-top: 4px solid var(--teal); }
.directory-member { display: grid; grid-template-columns: 150px minmax(220px, .72fr) minmax(320px, 1.28fr); gap: clamp(1.5rem, 3vw, 2.75rem); align-items: start; padding: clamp(1.5rem, 3vw, 2.25rem); background: var(--surface); border: 1px solid var(--line); border-top: 0; }
.directory-photo { width: 150px; aspect-ratio: 4 / 5; margin: 0; overflow: hidden; background: var(--ink); }
.directory-photo img { width: 100%; height: 100%; display: block; object-fit: cover; object-position: center top; }
.directory-monogram { width: 150px; aspect-ratio: 4 / 3; display: grid; place-items: center; color: white; background: var(--ink); font-size: 1.15rem; font-weight: 800; letter-spacing: .08em; }
.member-identity { padding-top: .2rem; }
.member-identity h2 { margin-bottom: .55rem; font-size: clamp(1.55rem, 2.2vw, 2.1rem); }
.member-title { margin-bottom: .3rem; color: var(--ink); font-size: .98rem; font-weight: 800; }
.member-affiliation { margin: 0; color: var(--muted); font-size: .84rem; }
.member-education { padding-left: clamp(1.25rem, 2.5vw, 2.25rem); border-left: 1px solid var(--line); }
.member-education h3 { margin-bottom: 1rem; color: var(--teal-dark); font-family: inherit; font-size: .72rem; font-weight: 800; letter-spacing: .1em; text-transform: uppercase; }
.education-list { display: grid; gap: .8rem; padding: 0; margin: 0; list-style: none; }
.education-list li { display: grid; grid-template-columns: minmax(0, .9fr) minmax(0, 1.1fr); gap: 1.25rem; padding-bottom: .7rem; border-bottom: 1px solid var(--line); }
.education-list li:last-child { padding-bottom: 0; border-bottom: 0; }
.education-list strong { color: var(--ink); font-size: .84rem; }
.education-list span { color: var(--muted); font-size: .82rem; }
.section-head-small { margin-bottom: 1.5rem; }
.section-head-small h2 { margin-bottom: 0; font-size: clamp(2rem, 3vw, 2.7rem); }
.directory-member-simple { grid-template-columns: 150px minmax(0, 1fr); align-items: center; }
.directory-member-simple .member-title { margin-bottom: 0; color: var(--muted); }

.timeline { position: relative; display: grid; gap: .75rem; }
.news-item { display: grid; grid-template-columns: 125px 1fr; gap: 2rem; padding: 1.65rem; background: var(--surface); border: 1px solid var(--line); border-left: 3px solid var(--teal); }
.news-date { color: var(--crimson); font-size: .86rem; font-weight: 800; letter-spacing: .045em; line-height: 1.4; text-transform: uppercase; }
.news-item h2 { font-size: 1.35rem; }
.news-item p { color: var(--muted); margin-bottom: 0; }
.grant-item { border-left-color: var(--crimson); }
.grant-source { margin-bottom: .55rem !important; color: var(--teal-dark) !important; font-size: .7rem; font-weight: 800; letter-spacing: .06em; text-transform: uppercase; }
.grant-details { display: flex; flex-wrap: wrap; gap: .65rem 1.5rem; padding: 1rem 0 0; margin: 1rem 0 0; border-top: 1px solid var(--line); list-style: none; }
.grant-details li { color: var(--ink); font-size: .8rem; font-weight: 700; }
.grant-details span { display: block; margin-bottom: .15rem; color: var(--muted); font-size: .6rem; font-weight: 800; letter-spacing: .09em; text-transform: uppercase; }

.contact-grid { display: grid; grid-template-columns: minmax(0, 760px); justify-content: start; }
.contact-card { padding: clamp(2rem, 4vw, 3.4rem); background: var(--surface); border: 1px solid var(--line); }
.contact-card.dark { color: white; background: var(--ink); border-color: var(--ink); border-top: 5px solid var(--teal); }
.contact-card h2 { font-size: 2rem; }
.contact-card.dark h2 { color: white; }
.contact-card p { color: var(--muted); }
.contact-card.dark p { color: #c3ccd5; }
.contact-card.dark .eyebrow { color: #9fd0cd; }
.contact-list { list-style: none; padding: 0; margin: 2rem 0 0; }
.contact-list li { padding: .9rem 0; border-top: 1px solid var(--line); }
.contact-card.dark .contact-list li { border-color: rgba(255,255,255,.16); }
.contact-list small { display: block; color: var(--muted); font-size: .65rem; font-weight: 800; letter-spacing: .11em; text-transform: uppercase; }
.contact-card.dark .contact-list small { color: #9da9b5; }

.site-footer { padding: 3rem 0; color: #adb8c4; background: var(--ink-deep); border-top: 4px solid var(--teal); }
.footer-grid { display: flex; align-items: end; justify-content: space-between; gap: 2rem; }
.footer-grid strong { display: block; color: white; margin-bottom: .3rem; font-size: 1rem; }
.footer-grid p { margin: 0; font-size: .8rem; }
.footer-links { display: flex; flex-wrap: wrap; gap: 1rem; }
.footer-links a { color: #cbd3db; font-size: .78rem; }

.reveal { opacity: 0; transform: translateY(10px); transition: opacity .45s ease, transform .45s ease; }
.reveal.visible { opacity: 1; transform: none; }

@media (min-width: 901px) {
  .nav-wrap { min-height: 68px; }
  .site-nav a { padding: 1.35rem 0 1.15rem; }
  .site-nav a::after { bottom: .72rem; }
  .hero-grid { height: clamp(480px, calc(100svh - 240px), 560px); min-height: 0; padding: 2.2rem 0; }
  .hero h1 { font-size: clamp(3rem, 5.1vw, 4.9rem); }
  .hero-lede { font-size: clamp(1rem, 1.35vw, 1.16rem); }
  .hero-actions { margin-top: 1.55rem; }
  .research-visual { min-height: 380px; height: min(390px, calc(100svh - 340px)); }
  .home-news { padding: 1rem 0 1.1rem; }
  .home-news-heading { margin-bottom: .6rem; }
  .home-news-heading .eyebrow { margin-bottom: .25rem; }
  .home-news-heading h2 { font-size: clamp(1.4rem, 2vw, 1.8rem); }
  .news-brief { min-height: 88px; padding: .85rem .95rem; }
}

@media (max-width: 900px) {
  .hero-grid, .section-head, .mission-grid, .page-hero .shell, .contact-grid { grid-template-columns: 1fr; }
  .hero-grid { padding: 6rem 0; gap: 3rem; }
  .research-visual { width: min(100%, 680px); }
  .news-strip { grid-template-columns: 1fr; }
  .news-brief { min-height: auto; border-right: 0; border-bottom: 1px solid var(--line); }
  .news-brief:last-child { border-bottom: 0; }
  .stats-grid { grid-template-columns: repeat(3, 1fr); padding: 1.4rem 0; }
  .stats-grid p { grid-column: 1 / span 2; }
  .research-explorer { grid-template-columns: 1fr; }
  .theme-tabs { grid-template-columns: repeat(3, 1fr); border-right: 0; border-bottom: 1px solid rgba(255,255,255,.16); }
  .theme-tab { border-right: 1px solid rgba(255,255,255,.13); border-bottom: 0; }
  .theme-tab:last-child { border-right: 0; }
  .theme-tab[aria-selected="true"] { box-shadow: inset 0 3px #65c4be; }
  .directory-member { grid-template-columns: 180px minmax(0, 1fr); }
  .directory-photo, .directory-monogram { width: 180px; }
  .member-education { grid-column: 2; padding: 1.5rem 0 0; border-top: 1px solid var(--line); border-left: 0; }
  .directory-member-simple { grid-template-columns: 180px minmax(0, 1fr); }
  .cards { grid-template-columns: 1fr 1fr; }
  .pipeline { grid-template-columns: 1fr 1fr; }
  .pipeline-step:nth-child(2) { border-right: 0; }
  .pipeline-step:nth-child(-n+2) { border-bottom: 1px solid var(--line); }
  .section-head { gap: 1rem; }
}

@media (max-width: 680px) {
  .shell { width: min(calc(100% - 28px), var(--max)); }
  .nav-wrap { min-height: 68px; }
  .nav-toggle { display: inline-grid; place-items: center; }
  .site-nav { position: absolute; left: 14px; right: 14px; top: 60px; display: none; padding: .5rem .9rem; border: 1px solid var(--line); background: var(--surface); box-shadow: var(--shadow); }
  .site-nav.open { display: grid; }
  .site-nav a { padding: .75rem 0; }
  .site-nav a::after { bottom: .45rem; right: auto; width: 28px; }
  .brand-text small { display: none; }
  .hero h1 { font-size: 3.25rem; }
  .hero-copy::before { display: none; }
  .research-visual { min-height: 410px; }
  .home-news-heading { align-items: flex-start; flex-direction: column; gap: .75rem; }
  .stats-grid { grid-template-columns: 1fr 1fr; gap: 1rem; }
  .stats-grid > div:nth-child(2) { border-right: 0; }
  .stats-grid > div:nth-child(3) { grid-column: 1 / -1; border-right: 0; }
  .stats-grid p, .stats-grid a { grid-column: 1 / -1; }
  .section { padding: 5rem 0; }
  .directory-member, .directory-member-simple { grid-template-columns: 1fr; gap: 1.25rem; }
  .directory-photo { width: 150px; }
  .directory-monogram { width: 150px; }
  .member-education { grid-column: auto; }
  .education-list li { grid-template-columns: 1fr; gap: .2rem; }
  .cards, .pipeline, .filters, .metric-grid { grid-template-columns: 1fr; }
  .theme-tabs { grid-template-columns: 1fr; }
  .theme-tab { border-right: 0; border-bottom: 1px solid rgba(255,255,255,.13); }
  .theme-tab:last-child { border-bottom: 0; }
  .theme-tab[aria-selected="true"] { box-shadow: inset 3px 0 #65c4be; }
  .theme-detail { padding: 2rem 1.35rem; }
  .evidence-grid { grid-template-columns: 1fr; }
  .evidence-grid > div { min-height: auto; }
  .pipeline-step, .pipeline-step:nth-child(2) { border-right: 0; border-bottom: 1px solid var(--line); }
  .pipeline-step:last-child { border-bottom: 0; }
  .publication, .news-item { grid-template-columns: 1fr; gap: .6rem; }
  .footer-grid { align-items: flex-start; flex-direction: column; }
}

@media (prefers-reduced-motion: reduce) {
  *, *::before, *::after { scroll-behavior: auto !important; animation: none !important; transition-duration: .01ms !important; }
  .reveal { opacity: 1; transform: none; }
}